🧬 NCERT Biology · Class 12 · Chapter 11
Organisms and Populations
Our living world is fascinatingly diverse and amazingly complex. We can try to understand its complexity by investigating processes at various levels of biological organisation–macromolecules, cells, tissues, organs, individual organisms, population, communities, ecosystems and biomes. At any level of biological organisation we can ask two types of questions – for example, when we hear the bulbul singing early morning in the garden, we may ask – ‘How does the bird sing?’ Or, ‘Why does the bird sing ?’ The ‘how-type’ questions seek the mechanism behind the process while the ‘why- type’ questions seek the significance of the process. For the first question in our example, the answer might be in terms of the operation of the voice box and the vibrating bone in the bird, whereas for the second question the answer may lie in the bird’s need to communicate with its mate during breeding season. When you observe nature around you with a scientific frame of mind you will certainly come up with many interesting questions of both types - Why are night-blooming flowers generally white? How does the bee know which flower has nectar? Why does cactus have so many thorns? How does the chick spures recognise her own mother ?, and so on.
You have already learnt in previous classes that Ecology is a subject which studies the interactions among organisms and between the organism and its physical (abiotic) environment. Ecology is basically concerned with four levels of biological organisation – organisms, populations, communities and biomes. In this chapter we explore ecology at population levels.
11.1 POPULATIONS
11.1.1 Population Attributes
In nature, we rarely find isolated, single individuals of any species; majority of them live in groups in a well defined geographical area, share or compete for similar resources, potentially interbreed and thus constitute a population. Although the term interbreeding implies sexual reproduction, a group of individuals resulting from even asexual reproduction is also generally considered a population for the purpose of ecological studies. All the cormorants in a wetland, rats in an abandoned dwelling, teakwood trees in a forest tract, bacteria in a culture plate and lotus plants in a pond, are some examples of a population. In earlier chapters you have learnt that although an individual organism is the one that has to cope with a changed environment, it is at the population level that natural selection operates to evolve the desired traits. Population ecology is, therefore, an important area because it links ecology to population genetics and evolution. A population has certain attributes whereas, an individual organism does not. An individual may have births and deaths, but a population has birth rates and death rates. In a population these rates refer to per capita births and deaths. The rates, hence, expressed are change in numbers (increase or decrease) with respect to members of the population. Here is an example. If in a pond there were 20 lotus plants last year and through reproduction 8 new plants are added, taking the current population to 28, we calculate the birth rate as 8/20 = 0.4 offspring per lotus per year. If 4 individuals in a laboratory population of 40 fruitflies died during a specified time interval, say a week, the death rate in the population during that period is 4/40 = 0.1 individuals per fruitfly per week. Another attribute characteristic of a population is sex ratio. An individual is either a male or a female but a population has a sex ratio (e.g., 60 per cent of the population are females and 40 per cent males). A population at any given time is composed of individuals of different ages. If the age distribution (per cent individuals of a given age or age group) is plotted for the population, the resulting structure is called an age pyramid (Figure 11.1). For human population, the age pyramids generally show age distribution of males and females in a diagram. The shape of the pyramids reflects the growth status of the population - (a) whether it is growing, (b) stable or (c) declining.
The size of the population tells us a lot about its status in the habitat. Whatever ecological processes we wish to investigate in a population, be it the outcome of competition with another species, the impact of a predator or the effect of a pesticide application, we always evaluate them in terms of any change in the population size. The size, in nature, could be as low as <10 (Siberian cranes at Bharatpur wetlands in any year) or go into millions (Chlamydomonas in a pond). Population size, technically called population density (designated as N), need not necessarily be measured in numbers only. Although total number is generally the most appropriate measure of population density, it is in some cases either meaningless or difficult to determine. In an area, if there are 200 carrot grass (Parthenium hysterophorus) plants but only a single huge banyan tree with a large canopy, stating that the population density of banyan is low relative to that of carrot grass amounts to underestimating the enormous role of the Banyan in that community. In such cases, the per cent cover or biomass is a more meaningful measure of the population size. Total number is again not an easily adoptable measure if the population is huge and counting is impossible or very time-consuming. If you have a dense laboratory culture of bacteria in a petri dish what is the best measure to report its density? Sometimes, for certain ecological investigations, there is no need to know the absolute population densities; relative densities serve the purpose equally well. For instance, the number of fish caught per trap is good enough measure of its total population density in the lake. We are mostly obliged to estimate population sizes indirectly, without actually counting them or seeing them. The tiger census in our national parks and tiger reserves is often based on pug marks and fecal pellets.
11.1.2 Population Growth
The size of a population for any species is not a static parameter. It keeps changing with time, depending on various factors including food availability, predation pressure and adverse weather. In fact, it is these changes in population density that give us some idea of what is happening to the population – whether it is flourishing or declining. Whatever might be the ultimate reasons, the density of a population in a given habitat during a given period, fluctuates due to changes in four basic processes,
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